"
WF <- createParam(cmd, overwrite = TRUE, writeParamFiles = TRUE, confirm = TRUE)
targetspath <- system.file("extdata", "targets.txt", package = "systemPipeR")
WF_test <- loadWorkflow(targets = targetspath, wf_file="hisat2.cwl",
input_file="hisat2.yml", dir_path = "param/cwl/hisat2/")
WF_test <- renderWF(WF_test, inputvars = c(FileName = "_FASTQ_PATH1_"))
WF_test
cmdlist(WF_test)[1:2]
```
# Inner Classes
`SYSargsList` steps are can be defined with two inner classes, `SYSargs2` and
`LineWise`. Next, more details on both classes.
## `SYSargs2` Class {#sysargs2}
*`SYSargs2`* workflow control class, an S4 class, is a list-like container where
each instance stores all the input/output paths and parameter components
required for a particular data analysis step. *`SYSargs2`* instances are
generated by two constructor functions, *loadWF* and *renderWF*, using as data
input *targets* or *yaml* files as well as two *cwl* parameter files (for
details see below).
In CWL, files with the extension *`.cwl`* define the parameters of a chosen
command-line step or workflow, while files with the extension *`.yml`* define
the input variables of command-line steps. Note, input variables provided by a
*targets* file can be passed on to a *`SYSargs2`* instance via the *inputvars*
argument of the *renderWF* function.
The following imports a *`.cwl`* file (here *`hisat2-mapping-se.cwl`*) for
running the short read aligner HISAT2 [@Kim2015-ve]. For more details about the
file structure and how to design or customize our own software tools, please
check `systemPipeR and CWL` pipeline.
```{r sysargs2_cwl_structure, echo = FALSE, eval=FALSE}
hisat2.cwl <- system.file("extdata", "cwl/hisat2/hisat2-mapping-se.cwl", package = "systemPipeR")
yaml::read_yaml(hisat2.cwl)
```
```{r sysargs2_yaml_structure, echo = FALSE, eval=FALSE}
hisat2.yml <- system.file("extdata", "cwl/hisat2/hisat2-mapping-se.yml", package = "systemPipeR")
yaml::read_yaml(hisat2.yml)
```
The *loadWF* and *renderWF* functions render the proper command-line strings for
each sample and software tool.
```{r SYSargs2_structure, eval=TRUE}
library(systemPipeR)
targetspath <- system.file("extdata", "targets.txt", package = "systemPipeR")
dir_path <- system.file("extdata/cwl", package = "systemPipeR")
WF <- loadWF(targets = targetspath, wf_file = "hisat2/hisat2-mapping-se.cwl",
input_file = "hisat2/hisat2-mapping-se.yml",
dir_path = dir_path)
WF <- renderWF(WF, inputvars = c(FileName = "_FASTQ_PATH1_",
SampleName = "_SampleName_"))
```
Several accessor methods are available that are named after the slot names of
the *`SYSargs2`* object.
```{r names_WF, eval=TRUE}
names(WF)
```
Of particular interest is the *`cmdlist()`* method. It constructs the system
commands for running command-line software as specified by a given *`.cwl`* file
combined with the paths to the input samples (*e.g.* FASTQ files) provided by a
*`targets`* file. The example below shows the *`cmdlist()`* output for running
HISAT2 on the first SE read sample. Evaluating the output of *`cmdlist()`* can
be very helpful for designing and debugging *`.cwl`* files of new command-line
software or changing the parameter settings of existing ones.
```{r cmdlist, eval=TRUE}
cmdlist(WF)[1]
```
The output components of *`SYSargs2`* define the expected output files for each
step in the workflow; some of which are the input for the next workflow step,
here next *`SYSargs2`* instance.
```{r output_WF, eval=TRUE}
output(WF)[1]
```
The targets components of `SYSargs2` object can be accessed by the targets
method. Here, for single-end (SE) samples, the structure of the targets file is
defined by:
- `FileName`: specify the FASTQ files path;
- `SampleName`: Unique IDs for each sample;
- `Factor`: ID for each treatment or condition.
```{r, targets_WF, eval=TRUE}
targets(WF)[1]
as(WF, "DataFrame")
```
Please note, to work with custom data, users need to generate a *`targets`* file
containing the paths to their own FASTQ files and then provide under
*`targetspath`* the path to the corresponding *`targets`* file.
In addition, if the [Environment Modules](http://modules.sourceforge.net/) is
available, it is possible to define which module should be loaded, as shown
here:
```{r, module_WF, eval=TRUE}
modules(WF)
```
Additional information can be accessed, as the parameters files location and the
`inputvars` provided to generate the object.
```{r, other_WF, eval=FALSE}
files(WF)
inputvars(WF)
```
## LineWise Class {#linewise}
`LineWise` was designed to store all the R code chunk when an RMarkdown file is
imported as a workflow.
```{r lw, eval=TRUE}
rmd <- system.file("extdata", "spr_simple_lw.Rmd", package = "systemPipeR")
sal_lw <- SPRproject(overwrite = TRUE)
sal_lw <- importWF(sal_lw, rmd, verbose = FALSE)
codeLine(sal_lw)
```
- Coerce methods available:
```{r, lw_coerce, eval=TRUE}
lw <- stepsWF(sal_lw)[[2]]
## Coerce
ll <- as(lw, "list")
class(ll)
lw <- as(ll, "LineWise")
lw
```
- Access details
```{r, lw_access, eval=TRUE}
length(lw)
names(lw)
codeLine(lw)
codeChunkStart(lw)
rmdPath(lw)
```
- Subsetting
```{r, lw_sub, eval=TRUE}
l <- lw[2]
codeLine(l)
l_sub <- lw[-2]
codeLine(l_sub)
```
- Replacement methods
```{r, lw_rep, eval=TRUE}
replaceCodeLine(lw, line = 2) <- "5+5"
codeLine(lw)
appendCodeLine(lw, after = 0) <- "6+7"
codeLine(lw)
```
- Replacement methods for `SYSargsList`
```{r, sal_rep_append, eval=FALSE}
replaceCodeLine(sal_lw, step = 2, line = 2) <- LineWise(code={
"5+5"
})
codeLine(sal_lw, step = 2)
appendCodeLine(sal_lw, step = 2) <- "66+55"
codeLine(sal_lw, step = 2)
appendCodeLine(sal_lw, step = 1, after = 1) <- "66+55"
codeLine(sal_lw, step = 1)
```
## Workflow design structure using *`SYSargs`*: Previous version
Instances of this S4 object class are constructed by the *`systemArgs`* function
from two simple tabular files: a *`targets`* file and a *`param`* file. The
latter is optional for workflow steps lacking command-line software. Typically,
a *`SYSargs`* instance stores all sample-level inputs as well as the paths to
the corresponding outputs generated by command-line- or R-based software
generating sample-level output files, such as read preprocessors
(trimmed/filtered FASTQ files), aligners (SAM/BAM files), variant callers
(VCF/BCF files) or peak callers (BED/WIG files). Each sample level input/output
operation uses its own *`SYSargs`* instance. The outpaths of *`SYSargs`* usually
define the sample inputs for the next *`SYSargs`* instance. This connectivity is
established by writing the outpaths with the *`writeTargetsout`* function to a
new *`targets`* file that serves as input to the next *`systemArgs`* call.
Typically, the user has to provide only the initial *`targets`* file. All
downstream *`targets`* files are generated automatically. By chaining several
*`SYSargs`* steps together one can construct complex workflows involving many
sample-level input/output file operations with any combination of command-line
or R-based software.
```{r, eval=TRUE, echo=FALSE, out.width="100%", fig.align = "center", fig.cap= "Workflow design structure of *`systemPipeR`* using previous version of *`SYSargs`*", warning=FALSE}
# knitr::include_graphics(system.file("extdata/images", "SystemPipeR_Workflow.png", package = "systemPipeR"))
```
# Third-party software tools {#third-party-software-tools}
Current, *systemPipeR* provides the _`param`_ file templates for third-party
software tools. Please check the listed software tools.
```{r table_tools, echo=FALSE, message=FALSE}
library(magrittr)
SPR_software <- system.file("extdata", "SPR_software.csv", package = "systemPipeR")
software <- read.delim(SPR_software, sep = ",", comment.char = "#")
colors <- colorRampPalette((c("darkseagreen", "indianred1")))(length(unique(software$Category)))
id <- as.numeric(c((unique(software$Category))))
software %>%
dplyr::mutate(Step = kableExtra::cell_spec(Step, color = "white", bold = TRUE,
background = factor(Category, id, colors)
)) %>%
dplyr::select(Tool, Description, Step) %>%
dplyr::arrange(Tool) %>%
kableExtra::kable(escape = FALSE, align = "c", col.names = c("Tool Name", "Description", "Step")) %>%
kableExtra::kable_styling(c("striped", "hover", "condensed"), full_width = TRUE) %>%
kableExtra::scroll_box(width = "80%", height = "500px")
```
Remember, if you desire to run any of these tools, make sure to have the
respective software installed on your system and configure in the `PATH`.
You can check as follows:
```{r test_tool_path, eval=FALSE}
tryCMD(command="gzip")
```
```{r cleaning3, eval=TRUE, include=FALSE}
if (file.exists(".SPRproject")) unlink(".SPRproject", recursive = TRUE)
## NOTE: Removing previous project create in the quick starts section
```
# Workflow commom steps overview
## Project initialization
To create a Workflow within _`systemPipeR`_, we can start by defining an empty
container and checking the directory structure:
```{r SPRproject2, eval=FALSE}
sal <- SPRproject()
```
### Required packages and resources
The `systemPipeR` package needs to be loaded [@H_Backman2016-bt].
```{r load_SPR, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
library(systemPipeR)
},
step_name = "load_SPR")
```
## Read Preprocessing
### Preprocessing with `preprocessReads` function
The function `preprocessReads` allows to apply predefined or custom
read preprocessing functions to all FASTQ files referenced in a
`SYSargsList` container, such as quality filtering or adapter trimming
routines. Internally, `preprocessReads` uses the `FastqStreamer` function from
the `ShortRead` package to stream through large FASTQ files in a
memory-efficient manner. The following example performs adapter trimming with
the `trimLRPatterns` function from the `Biostrings` package.
Here, we are appending this step to the `SYSargsList` object created previously.
All the parameters are defined on the `preprocessReads/preprocessReads-pe.yml` file.
```{r preprocessing, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(
step_name = "preprocessing",
targets = "targetsPE.txt", dir = TRUE,
wf_file = "preprocessReads/preprocessReads-pe.cwl",
input_file = "preprocessReads/preprocessReads-pe.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = c(
FileName1 = "_FASTQ_PATH1_",
FileName2 = "_FASTQ_PATH2_",
SampleName = "_SampleName_"
),
dependency = c("load_SPR"))
```
After the preprocessing step, the `outfiles` files can be used to generate the new
targets files containing the paths to the trimmed FASTQ files. The new targets
information can be used for the next workflow step instance, _e.g._ running the
NGS alignments with the trimmed FASTQ files. The `appendStep` function is
automatically handling this connectivity between steps. Please check the next
step for more details.
The following example shows how one can design a custom read _'preprocessReads'_
function using utilities provided by the `ShortRead` package, and then run it
in batch mode with the _'preprocessReads'_ function. Here, it is possible to
replace the function used on the `preprocessing` step and modify the `sal` object.
Because it is a custom function, it is necessary to save the part in the R object,
and internally the `preprocessReads.doc.R` is loading the custom function.
If the R object is saved with a different name (here `"param/customFCT.RData"`),
please replace that accordingly in the `preprocessReads.doc.R`.
Please, note that this step is not added to the workflow, here just for
demonstration.
First, we defined the custom function in the workflow:
```{r custom_preprocessing_function, eval=FALSE}
appendStep(sal) <- LineWise(
code = {
filterFct <- function(fq, cutoff = 20, Nexceptions = 0) {
qcount <- rowSums(as(quality(fq), "matrix") <= cutoff, na.rm = TRUE)
# Retains reads where Phred scores are >= cutoff with N exceptions
fq[qcount <= Nexceptions]
}
save(list = ls(), file = "param/customFCT.RData")
},
step_name = "custom_preprocessing_function",
dependency = "preprocessing"
)
```
After, we can edit the input parameter:
```{r editing_preprocessing, message=FALSE, eval=FALSE}
yamlinput(sal, "preprocessing")$Fct
yamlinput(sal, "preprocessing", "Fct") <- "'filterFct(fq, cutoff=20, Nexceptions=0)'"
yamlinput(sal, "preprocessing")$Fct ## check the new function
cmdlist(sal, "preprocessing", targets = 1) ## check if the command line was updated with success
```
### Preprocessing with TrimGalore!
[TrimGalore!](http://www.bioinformatics.babraham.ac.uk/projects/trim_galore/) is
a wrapper tool to consistently apply quality and adapter trimming to fastq files,
with some extra functionality for removing Reduced Representation Bisulfite-Seq
(RRBS) libraries.
```{r trimGalore, eval=FALSE, spr=TRUE}
targetspath <- system.file("extdata", "targets.txt", package = "systemPipeR")
appendStep(sal) <- SYSargsList(step_name = "trimGalore",
targets = targetspath, dir = TRUE,
wf_file = "trim_galore/trim_galore-se.cwl",
input_file = "trim_galore/trim_galore-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = c(FileName = "_FASTQ_PATH1_", SampleName = "_SampleName_"),
dependency = "load_SPR",
run_step = "optional")
```
### Preprocessing with Trimmomatic
[Trimmomatic](http://www.usadellab.org/cms/?page=trimmomatic) software [@Bolger2014-yr]
performs a variety of useful trimming tasks for Illumina paired-end and single
ended data. Here, an example of how to perform this task using parameters template
files for trimming FASTQ files.
```{r trimmomatic, eval=FALSE, spr=TRUE}
targetspath <- system.file("extdata", "targets.txt", package = "systemPipeR")
appendStep(sal) <- SYSargsList(step_name = "trimmomatic",
targets = targetspath, dir = TRUE,
wf_file = "trimmomatic/trimmomatic-se.cwl",
input_file = "trimmomatic/trimmomatic-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = c(FileName = "_FASTQ_PATH1_", SampleName = "_SampleName_"),
dependency = "load_SPR",
run_step = "optional")
```
## FASTQ quality report
The following `seeFastq` and `seeFastqPlot` functions generate and plot a series of useful
quality statistics for a set of FASTQ files, including per cycle quality box
plots, base proportions, base-level quality trends, relative k-mer
diversity, length, and occurrence distribution of reads, number of reads
above quality cutoffs and mean quality distribution. The results are
written to a PDF file named `fastqReport.pdf`.
```{r fastq_report, eval=FALSE, message=FALSE, spr=TRUE}
appendStep(sal) <- LineWise(code = {
fastq <- getColumn(sal, step = "preprocessing", "targetsWF", column = 1)
fqlist <- seeFastq(fastq = fastq, batchsize = 10000, klength = 8)
pdf("./results/fastqReport.pdf", height = 18, width = 4*length(fqlist))
seeFastqPlot(fqlist)
dev.off()
}, step_name = "fastq_report",
dependency = "preprocessing")
```
**Figure 1:** FASTQ quality report
## NGS Alignment software
After quality control, the sequence reads can be aligned to a reference genome or
transcriptome database. The following sessions present some NGS sequence alignment
software. Select the most accurate aligner and determining the optimal parameter
for your custom data set project.
For all the following examples, it is necessary to install the respective software
and export the `PATH` accordingly.
### Alignment with `HISAT2`
The following steps will demonstrate how to use the short read aligner `Hisat2`
[@Kim2015-ve] in both interactive job submissions and batch submissions to
queuing systems of clusters using the _`systemPipeR's`_ new CWL command-line interface.
- Build `Hisat2` index.
```{r hisat_index, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "hisat_index",
targets = NULL, dir = FALSE,
wf_file = "hisat2/hisat2-index.cwl",
input_file = "hisat2/hisat2-index.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = NULL,
dependency = "preprocessing")
```
The parameter settings of the aligner are defined in the `workflow_hisat2-se.cwl`
and `workflow_hisat2-se.yml` files. The following shows how to construct the
corresponding *SYSargsList* object, and append to *sal* workflow.
- Alignment with `HISAT2` and `SAMtools`
It possible to build an workflow with `HISAT2` and `SAMtools`.
```{r hisat_mapping_samtools, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "hisat_mapping",
targets = "preprocessing", dir = TRUE,
wf_file = "workflow-hisat2/workflow_hisat2-se.cwl",
input_file = "workflow-hisat2/workflow_hisat2-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars=c(preprocessReads_se="_FASTQ_PATH1_", SampleName="_SampleName_"),
dependency = c("hisat_index"),
run_session = "remote")
```
### Alignment with `Tophat2`
The NGS reads of this project can also be aligned against the reference genome
sequence using `Bowtie2/TopHat2` [@Kim2013-vg; @Langmead2012-bs].
- Build _`Bowtie2`_ index.
```{r bowtie_index, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "bowtie_index",
targets = NULL, dir = FALSE,
wf_file = "bowtie2/bowtie2-index.cwl",
input_file = "bowtie2/bowtie2-index.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = NULL,
dependency = "preprocessing",
run_step = "optional")
```
The parameter settings of the aligner are defined in the `workflow_tophat2-mapping.cwl`
and `tophat2-mapping-pe.yml` files. The following shows how to construct the
corresponding *SYSargsList* object, using the outfiles from the `preprocessing` step.
```{r tophat2_mapping, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "tophat2_mapping",
targets = "preprocessing", dir = TRUE,
wf_file = "tophat2/workflow_tophat2-mapping-se.cwl",
input_file = "tophat2/tophat2-mapping-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars=c(preprocessReads_se="_FASTQ_PATH1_", SampleName="_SampleName_"),
dependency = c("bowtie_index"),
run_session = "remote",
run_step = "optional")
```
### Alignment with _`Bowtie2`_ (_e.g._ for miRNA profiling)
The following example runs _`Bowtie2`_ as a single process without submitting it to a cluster.
```{r bowtie2_mapping, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "bowtie2_mapping",
targets = "preprocessing", dir = TRUE,
wf_file = "bowtie2/workflow_bowtie2-mapping-se.cwl",
input_file = "bowtie2/bowtie2-mapping-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars=c(preprocessReads_se="_FASTQ_PATH1_", SampleName="_SampleName_"),
dependency = c("bowtie_index"),
run_session = "remote",
run_step = "optional")
```
### Alignment with _`BWA-MEM`_ (_e.g._ for VAR-Seq)
The following example runs BWA-MEM as a single process without submitting it to a cluster.
- Build the index:
```{r bwa_index, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "bwa_index",
targets = NULL, dir = FALSE,
wf_file = "bwa/bwa-index.cwl",
input_file = "bwa/bwa-index.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = NULL,
dependency = "preprocessing",
run_step = "optional")
```
- Prepare the alignment step:
```{r bwa_mapping, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "bwa_mapping",
targets = "preprocessing", dir = TRUE,
wf_file = "bwa/bwa-se.cwl",
input_file = "bwa/bwa-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars=c(preprocessReads_se="_FASTQ_PATH1_", SampleName="_SampleName_"),
dependency = c("bwa_index"),
run_session = "remote",
run_step = "optional")
```
### Alignment with _`Rsubread`_ (_e.g._ for RNA-Seq)
The following example shows how one can use within the \Rpackage{systemPipeR} environment the R-based aligner \Rpackage{Rsubread}, allowing running from R or command-line.
- Build the index:
```{r rsubread_index, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "rsubread_index",
targets = NULL, dir = FALSE,
wf_file = "rsubread/rsubread-index.cwl",
input_file = "rsubread/rsubread-index.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = NULL,
dependency = "preprocessing",
run_step = "optional")
```
- Prepare the alignment step:
```{r rsubread_mapping, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "rsubread",
targets = "preprocessing", dir = TRUE,
wf_file = "rsubread/rsubread-mapping-se.cwl",
input_file = "rsubread/rsubread-mapping-se.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars=c(preprocessReads_se="_FASTQ_PATH1_", SampleName="_SampleName_"),
dependency = c("rsubread_index"),
run_session = "remote",
run_step = "optional")
```
### Alignment with _`gsnap`_ (_e.g._ for VAR-Seq and RNA-Seq)
Another R-based short read aligner is _`gsnap`_ from the _`gmapR`_ package [@Wu2010-iq].
The code sample below introduces how to run this aligner on multiple nodes of a compute cluster.
- Build the index:
```{r gsnap_index, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "gsnap_index",
targets = NULL, dir = FALSE,
wf_file = "gsnap/gsnap-index.cwl",
input_file = "gsnap/gsnap-index.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = NULL,
dependency = "preprocessing",
run_step = "optional")
```
- Prepare the alignment step:
```{r gsnap_mapping, eval=FALSE, spr=TRUE}
appendStep(sal) <- SYSargsList(step_name = "gsnap",
targets = "targetsPE.txt", dir = TRUE,
wf_file = "gsnap/gsnap-mapping-pe.cwl",
input_file = "gsnap/gsnap-mapping-pe.yml",
dir_path = system.file("extdata/cwl", package = "systemPipeR"),
inputvars = c(FileName1 = "_FASTQ_PATH1_",
FileName2 = "_FASTQ_PATH2_", SampleName = "_SampleName_"),
dependency = c("gsnap_index"),
run_session = "remote",
run_step = "optional")
```
## Create symbolic links for viewing BAM files in IGV
The genome browser IGV supports reading of indexed/sorted BAM files via web URLs. This way it can be avoided to create unnecessary copies of these large files. To enable this approach, an HTML directory with Http access needs to be available in the user account (_e.g._ _`home/publichtml`_) of a system. If this is not the case then the BAM files need to be moved or copied to the system where IGV runs. In the following, _`htmldir`_ defines the path to the HTML directory with http access where the symbolic links to the BAM files will be stored. The corresponding URLs will be written to a text file specified under the `_urlfile`_ argument.
```{r bam_IGV, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise(
code = {
bampaths <- getColumn(sal, step = "hisat2_mapping", "outfiles",
column = "samtools_sort_bam")
symLink2bam(
sysargs = bampaths, htmldir = c("~/.html/", "somedir/"),
urlbase = "http://cluster.hpcc.ucr.edu/~tgirke/",
urlfile = "./results/IGVurl.txt")
},
step_name = "bam_IGV",
dependency = "hisat2_mapping",
run_step = "optional"
)
```
## Read counting for mRNA profiling experiments
Create _`txdb`_ (needs to be done only once).
```{r create_txdb, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise(code = {
library(GenomicFeatures)
txdb <- makeTxDbFromGFF(file="data/tair10.gff", format="gff",
dataSource="TAIR", organism="Arabidopsis thaliana")
saveDb(txdb, file="./data/tair10.sqlite")
},
step_name = "create_txdb",
dependency = "hisat_mapping")
```
The following performs read counting with _`summarizeOverlaps`_ in parallel mode with multiple cores.
```{r read_counting, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
library(BiocParallel)
txdb <- loadDb("./data/tair10.sqlite")
eByg <- exonsBy(txdb, by="gene")
outpaths <- getColumn(sal, step = "hisat_mapping", 'outfiles', column = 2)
bfl <- BamFileList(outpaths, yieldSize=50000, index=character())
multicoreParam <- MulticoreParam(workers=4); register(multicoreParam); registered()
counteByg <- bplapply(bfl, function(x) summarizeOverlaps(eByg, x, mode="Union",
ignore.strand=TRUE,
inter.feature=TRUE,
singleEnd=TRUE))
# Note: for strand-specific RNA-Seq set 'ignore.strand=FALSE' and for PE data set 'singleEnd=FALSE'
countDFeByg <- sapply(seq(along=counteByg),
function(x) assays(counteByg[[x]])$counts)
rownames(countDFeByg) <- names(rowRanges(counteByg[[1]]))
colnames(countDFeByg) <- names(bfl)
rpkmDFeByg <- apply(countDFeByg, 2, function(x) returnRPKM(counts=x, ranges=eByg))
write.table(countDFeByg, "results/countDFeByg.xls",
col.names=NA, quote=FALSE, sep="\t")
write.table(rpkmDFeByg, "results/rpkmDFeByg.xls",
col.names=NA, quote=FALSE, sep="\t")
},
step_name = "read_counting",
dependency = "create_txdb")
```
Please note, in addition to read counts this step generates RPKM normalized expression values. For most statistical differential expression or abundance analysis methods, such as _`edgeR`_ or _`DESeq2`_, the raw count values should be used as input. The usage of RPKM values should be restricted to specialty applications required by some users, _e.g._ manually comparing the expression levels of different genes or features.
#### Read and alignment count stats
Generate a table of read and alignment counts for all samples.
```{r align_stats, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
read_statsDF <- alignStats(args)
write.table(read_statsDF, "results/alignStats.xls",
row.names = FALSE, quote = FALSE, sep = "\t")
},
step_name = "align_stats",
dependency = "hisat_mapping")
```
The following shows the first four lines of the sample alignment stats file
provided by the _`systemPipeR`_ package. For simplicity the number of PE reads
is multiplied here by 2 to approximate proper alignment frequencies where each
read in a pair is counted.
```{r align_stats2, eval=TRUE}
read.table(system.file("extdata", "alignStats.xls", package = "systemPipeR"), header = TRUE)[1:4,]
```
## Read counting for miRNA profiling experiments
Download `miRNA` genes from `miRBase`.
```{r read_counting_mirna, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
system("wget https://www.mirbase.org/ftp/CURRENT/genomes/ath.gff3 -P ./data/")
gff <- rtracklayer::import.gff("./data/ath.gff3")
gff <- split(gff, elementMetadata(gff)$ID)
bams <- getColumn(sal, step = "bowtie2_mapping", 'outfiles', column = 2)
bfl <- BamFileList(bams, yieldSize=50000, index=character())
countDFmiR <- summarizeOverlaps(gff, bfl, mode="Union",
ignore.strand = FALSE, inter.feature = FALSE)
countDFmiR <- assays(countDFmiR)$counts
# Note: inter.feature=FALSE important since pre and mature miRNA ranges overlap
rpkmDFmiR <- apply(countDFmiR, 2, function(x) returnRPKM(counts = x, ranges = gff))
write.table(assays(countDFmiR)$counts, "results/countDFmiR.xls",
col.names=NA, quote=FALSE, sep="\t")
write.table(rpkmDFmiR, "results/rpkmDFmiR.xls", col.names=NA, quote=FALSE, sep="\t")
},
step_name = "read_counting_mirna",
dependency = "bowtie2_mapping")
```
## Correlation analysis of samples
The following computes the sample-wise Spearman correlation coefficients from the _`rlog`_ (regularized-logarithm) transformed expression values generated with the _`DESeq2`_ package. After transformation to a distance matrix, hierarchical clustering is performed with the _`hclust`_ function and the result is plotted as a dendrogram ([sample\_tree.pdf](sample_tree.png)).
```{r sample_tree_rlog, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
library(DESeq2, warn.conflicts=FALSE, quietly=TRUE)
library(ape, warn.conflicts=FALSE)
countDFpath <- system.file("extdata", "countDFeByg.xls", package="systemPipeR")
countDF <- as.matrix(read.table(countDFpath))
colData <- data.frame(row.names = targetsWF(sal)[[2]]$SampleName,
condition=targetsWF(sal)[[2]]$Factor)
dds <- DESeqDataSetFromMatrix(countData = countDF, colData = colData,
design = ~ condition)
d <- cor(assay(rlog(dds)), method = "spearman")
hc <- hclust(dist(1-d))
plot.phylo(as.phylo(hc), type = "p", edge.col = 4, edge.width = 3,
show.node.label = TRUE, no.margin = TRUE)
},
step_name = "sample_tree_rlog",
dependency = "read_counting")
```
**Figure 2:** Correlation dendrogram of samples for _`rlog`_ values.
## DEG analysis with _`edgeR`_
The following _`run_edgeR`_ function is a convenience wrapper for
identifying differentially expressed genes (DEGs) in batch mode with
_`edgeR`_'s GML method [@Robinson2010-uk] for any number of
pairwise sample comparisons specified under the _`cmp`_ argument. Users
are strongly encouraged to consult the
[_`edgeR`_](\href{http://www.bioconductor.org/packages/devel/bioc/vignettes/edgeR/inst/doc/edgeRUsersGuide.pdf) vignette
for more detailed information on this topic and how to properly run _`edgeR`_
on data sets with more complex experimental designs.
```{r edger, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
targetspath <- system.file("extdata", "targets.txt", package = "systemPipeR")
targets <- read.delim(targetspath, comment = "#")
cmp <- readComp(file = targetspath, format = "matrix", delim = "-")
countDFeBygpath <- system.file("extdata", "countDFeByg.xls", package = "systemPipeR")
countDFeByg <- read.delim(countDFeBygpath, row.names = 1)
edgeDF <- run_edgeR(countDF = countDFeByg, targets = targets, cmp = cmp[[1]],
independent = FALSE, mdsplot = "")
DEG_list <- filterDEGs(degDF = edgeDF, filter = c(Fold = 2, FDR = 10))
},
step_name = "edger",
dependency = "read_counting")
```
Filter and plot DEG results for up and down-regulated genes. Because of the small size of the toy data set used by this vignette, the _FDR_ value has been set to a relatively high threshold (here 10%). More commonly used _FDR_ cutoffs are 1% or 5%. The definition of '_up_' and '_down_' is given in the corresponding help file. To open it, type _`?filterDEGs`_ in the R console.
**Figure 3:** Up and down regulated DEGs identified by _`edgeR`_.
## DEG analysis with _`DESeq2`_
The following _`run_DESeq2`_ function is a convenience wrapper for
identifying DEGs in batch mode with _`DESeq2`_ [@Love2014-sh] for any number of
pairwise sample comparisons specified under the _`cmp`_ argument. Users
are strongly encouraged to consult the
[_`DESeq2`_](http://www.bioconductor.org/packages/devel/bioc/vignettes/DESeq2/inst/doc/DESeq2.pdf) vignette
for more detailed information on this topic and how to properly run _`DESeq2`_
on data sets with more complex experimental designs.
```{r deseq2, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
degseqDF <- run_DESeq2(countDF=countDFeByg, targets=targets, cmp=cmp[[1]],
independent=FALSE)
DEG_list2 <- filterDEGs(degDF=degseqDF, filter=c(Fold=2, FDR=10))
},
step_name = "deseq2",
dependency = "read_counting")
```
## Venn Diagrams
The function _`overLapper`_ can compute Venn intersects for large numbers of sample sets (up to 20 or more) and _`vennPlot`_ can plot 2-5 way Venn diagrams. A useful feature is the possibility to combine the counts from several Venn comparisons with the same number of sample sets in a single Venn diagram (here for 4 up and down DEG sets).
```{r vennplot, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
vennsetup <- overLapper(DEG_list$Up[6:9], type="vennsets")
vennsetdown <- overLapper(DEG_list$Down[6:9], type="vennsets")
vennPlot(list(vennsetup, vennsetdown), mymain="", mysub="",
colmode=2, ccol=c("blue", "red"))
},
step_name = "vennplot",
dependency = "edger")
```
**Figure 4:** Venn Diagram for 4 Up and Down DEG Sets.
## GO term enrichment analysis of DEGs
### Obtain gene-to-GO mappings
The following shows how to obtain gene-to-GO mappings from _`biomaRt`_ (here for _A. thaliana_) and how to organize them for the downstream GO term enrichment analysis. Alternatively, the gene-to-GO mappings can be obtained for many organisms from Bioconductor's _`*.db`_ genome annotation packages or GO annotation files provided by various genome databases. For each annotation, this relatively slow preprocessing step needs to be performed only once. Subsequently, the preprocessed data can be loaded with the _`load`_ function as shown in the next subsection.
```{r get_go_biomart, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
library("biomaRt")
listMarts() # To choose BioMart database
listMarts(host="plants.ensembl.org")
m <- useMart("plants_mart", host="https://plants.ensembl.org")
listDatasets(m)
m <- useMart("plants_mart", dataset="athaliana_eg_gene", host="https://plants.ensembl.org")
listAttributes(m) # Choose data types you want to download
go <- getBM(attributes=c("go_id", "tair_locus", "namespace_1003"), mart=m)
go <- go[go[,3]!="",]; go[,3] <- as.character(go[,3])
go[go[,3]=="molecular_function", 3] <- "F"
go[go[,3]=="biological_process", 3] <- "P"
go[go[,3]=="cellular_component", 3] <- "C"
go[1:4,]
dir.create("./data/GO")
write.table(go, "data/GO/GOannotationsBiomart_mod.txt",
quote=FALSE, row.names=FALSE, col.names=FALSE, sep="\t")
catdb <- makeCATdb(myfile="data/GO/GOannotationsBiomart_mod.txt",
lib=NULL, org="", colno=c(1,2,3), idconv=NULL)
save(catdb, file="data/GO/catdb.RData")
},
step_name = "get_go_biomart",
dependency = "edger")
```
### Batch GO term enrichment analysis
Apply the enrichment analysis to the DEG sets obtained in the above differential expression analysis. Note, in the following example the _FDR_ filter is set here to an unreasonably high value, simply because of the small size of the toy data set used in this vignette. Batch enrichment analysis of many gene sets is performed with the _`GOCluster_Report`_ function. When _`method="all"`_, it returns all GO terms passing the p-value cutoff specified under the _`cutoff`_ arguments. When _`method="slim"`_, it returns only the GO terms specified under the _`myslimv`_ argument. The given example shows how one can obtain such a GO slim vector from BioMart for a specific organism.
```{r go_enrichment, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
load("data/GO/catdb.RData")
DEG_list <- filterDEGs(degDF=edgeDF, filter=c(Fold=2, FDR=50), plot=FALSE)
up_down <- DEG_list$UporDown; names(up_down) <- paste(names(up_down), "_up_down", sep="")
up <- DEG_list$Up; names(up) <- paste(names(up), "_up", sep="")
down <- DEG_list$Down; names(down) <- paste(names(down), "_down", sep="")
DEGlist <- c(up_down, up, down)
DEGlist <- DEGlist[sapply(DEGlist, length) > 0]
BatchResult <- GOCluster_Report(catdb=catdb, setlist=DEGlist, method="all",
id_type="gene", CLSZ=2, cutoff=0.9,
gocats=c("MF", "BP", "CC"), recordSpecGO=NULL)
library("biomaRt")
m <- useMart("plants_mart", dataset="athaliana_eg_gene", host="https://plants.ensembl.org")
goslimvec <- as.character(getBM(attributes=c("goslim_goa_accession"), mart=m)[,1])
BatchResultslim <- GOCluster_Report(catdb=catdb, setlist=DEGlist, method="slim",
id_type="gene", myslimv=goslimvec, CLSZ=10,
cutoff=0.01, gocats=c("MF", "BP", "CC"),
recordSpecGO=NULL)
gos <- BatchResultslim[grep("M6-V6_up_down", BatchResultslim$CLID), ]
gos <- BatchResultslim
pdf("GOslimbarplotMF.pdf", height=8, width=10); goBarplot(gos, gocat="MF"); dev.off()
goBarplot(gos, gocat="BP")
goBarplot(gos, gocat="CC")
},
step_name = "go_enrichment",
dependency = "get_go_biomart")
```
### Plot batch GO term results
The _`data.frame`_ generated by _`GOCluster_Report`_ can be plotted with the _`goBarplot`_ function. Because of the variable size of the sample sets, it may not always be desirable to show the results from different DEG sets in the same bar plot. Plotting single sample sets is achieved by subsetting the input data frame as shown in the first line of the following example.
**Figure 5:** GO Slim Barplot for MF Ontology.
## Clustering and heat maps
The following example performs hierarchical clustering on the _`rlog`_ transformed expression matrix subsetted by the DEGs identified in the
above differential expression analysis. It uses a Pearson correlation-based distance measure and complete linkage for cluster join.
```{r hierarchical_clustering, message=FALSE, eval=FALSE, spr=TRUE}
appendStep(sal) <- LineWise({
library(pheatmap)
geneids <- unique(as.character(unlist(DEG_list[[1]])))
y <- assay(rlog(dds))[geneids, ]
pdf("heatmap1.pdf")
pheatmap(y, scale="row", clustering_distance_rows="correlation",
clustering_distance_cols="correlation")
dev.off()
},
step_name = "hierarchical_clustering",
dependency = c("sample_tree_rlog", "edgeR"))
```
**Figure 7:** Heat map with hierarchical clustering dendrograms of DEGs.
## Visualize workflow
systemPipeR workflows instances can be visualized with the `plotWF` function.
This function will make a plot of selected workflow instance and the following information is displayed on the plot:
- Workflow structure (dependency graphs between different steps);
- Workflow step status, *e.g.* `Success`, `Error`, `Pending`, `Warnings`;
- Sample status and statistics;
- Workflow timing: running duration time.
If no argument is provided, the basic plot will automatically detect width, height, layout, plot method, branches, etc.
```{r}
plotWF(sal, show_legend = TRUE, width = "80%")
```
To check more details of `plotWF`, visit [our website](https://systempipe.org/sp/spr/sp_run/step_vis/).
## Running workflow
### Interactive job submissions in a single machine
For running the workflow, `runWF` function will execute all the steps store in
the workflow container. The execution will be on a single machine without
submitting to a queuing system of a computer cluster.
```{r runWF_steps, eval=FALSE}
sal <- runWF(sal)
```
### Parallelization on clusters
Alternatively, the computation can be greatly accelerated by processing many files
in parallel using several compute nodes of a cluster, where a scheduling/queuing
system is used for load balancing.
The `resources` list object provides the number of independent parallel cluster
processes defined under the `Njobs` element in the list. The following example
will run 18 processes in parallel using each 4 CPU cores.
If the resources available on a cluster allow running all 18 processes at the
same time, then the shown sample submission will utilize in a total of 72 CPU cores.
Note, `runWF` can be used with most queueing systems as it is based on utilities
from the `batchtools` package, which supports the use of template files (_`*.tmpl`_)
for defining the run parameters of different schedulers. To run the following
code, one needs to have both a `conffile` (see _`.batchtools.conf.R`_ samples [here](https://mllg.github.io/batchtools/))
and a `template` file (see _`*.tmpl`_ samples [here](https://github.com/mllg/batchtools/tree/master/inst/templates))
for the queueing available on a system. The following example uses the sample
`conffile` and `template` files for the Slurm scheduler provided by this package.
The resources can be appended when the step is generated, or it is possible to
add these resources later, as the following example using the `addResources`
function:
```{r runWF_cluster_steps, eval=FALSE}
resources <- list(conffile=".batchtools.conf.R",
template="batchtools.slurm.tmpl",
Njobs=18,
walltime=120, ## minutes
ntasks=1,
ncpus=4,
memory=1024, ## Mb
partition = "short"
)
sal <- addResources(sal, c("hisat2_mapping"), resources = resources)
sal <- runWF(sal)
```
### Checking workflow status
To check the summary of the workflow, we can use:
```{r statusWF_steps, eval=FALSE}
sal
statusWF(sal)
```
### Accessing logs report
_`systemPipeR`_ compiles all the workflow execution logs in one central location,
making it easier to check any standard output (`stdout`) or standard error
(`stderr`) for any command-line tools used on the workflow or the R code stdout.
```{r logsWF_steps, eval=FALSE}
sal <- renderLogs(sal)
```
```{r quiting, eval=TRUE, echo=FALSE}
knitr::opts_knit$set(root.dir = '../')
unlink("rnaseq", recursive = TRUE)
```
# Version information
```{r sessionInfo}
sessionInfo()
```
# Funding
This project is funded by NSF award [ABI-1661152](https://www.nsf.gov/awardsearch/showAward?AWD_ID=1661152).
# References